2002: 华东师范大学化学系，学士；一等奖学金
2007: 华东师范大学河口海岸学国家重点实验室，博士;上海市优秀毕业生
2005: 德国极地与海洋研究所，访问博士生

海洋有机地球化学
海洋氧亏损（低氧）的生消机制及其对碳氮循环的影响和效应
海洋有机碳的通量、示踪和命运

在当前的一段时间内，我在开展海洋和河流、河口环境中（包括北极、南极、赤道海洋；以及不同国家内不同生境的河口近海）有机质性质与成分的研究：通过生物标志物与其他环境参数的分析，揭示有机化合物在海洋、河流中的分布特点，探索有机质的成分变化规律，诠释其背后的控制因素；探索海洋作为有机碳库储存人类排放的过剩二氧化碳的过程中的不确定性，帮助解释当前人类强烈活动背景下全球气候和海洋变化所面临的挑战。
另外，我对水体中的溶解氧也很感兴趣——事实上在我看来这是有机质循环的一部分：有机质在表层经由初级生产产生，这伴随着溶解氧的增加；有机质在底层水的降解，则伴随着溶解氧的亏损。因此，对溶解氧的研究自然就包括其含量、变化特点和控制因素，以及背后的故事。例如，我对长江口外的低氧问题开展了多年的追踪。通过接近10次的船载现场调查，和大量室内仪器分析，我和我的同事们在过去的10年里跟踪了长江口外低氧区的面积和变化规律、并就南北两个区域的氧亏损-有机质降解-营养盐再生的耦合机理差异进行了定量表达。同时，我还在船上进行培养实验，定量现场体系的呼吸耗氧速率，以帮助同事建立低氧模型，为今后预测低氧的发生，乃至低氧的治理提供科学基础。最近两年以来，通过溶解氧同位素技术，我和我的同事发现水柱呼吸和沉积物耗氧对低氧的贡献比例很可能并非一成不变。在低氧发生的时候，水柱呼吸过程（沉降颗粒物的降解）是底层水缺氧的核心推手，海底界面的沉积物耗氧对低氧发生贡献是次要的。于我，这是一个激动人心的定量发现。
在之后的继续研究中，我又把这个假设推广到了潮滩等极浅水体系，发现上述沉积物和水柱耗氧的演替规律仍旧适用。这在氧亏损机制的前进的道路上给予了我新的信心。目前我聚焦于解构混合过程对SOR干扰的问题。

基于上述描述，你可以叫我海洋学家或海洋学工作者。用学科的术语来说，我的专业是海洋化学，研究兴趣是溶解氧亏损生消机制及其有机碳循环效应。
欢迎对低氧（氧亏损）、碳氮循环感兴趣的同学加入OMEGA小组，读硕读博、博士后均欢迎。

2007-2010: 华东师范大学生物系,博士后
2008.12，2009.12：德国极地与海洋研究所，访问学者
2010-2013: 华东师范大学河口海岸学国家重点实验室；助理研究员
2011.3：印度国家海洋研究所，访问学者
2014-2020：华东师范大学河口海岸学国家重点实验室；副研究员，博导
2013.12-2014.3：华盛顿大学（西雅图），访问学者
2015.4-5，2016.5-6：普林斯顿大学，访问学者
2020-2023：上海交通大学海洋学院；副研究员，博导
2024-：上海交通大学海洋学院；研究员，博导
2024-：中国极地研究中心（中国极地研究所）双聘研究员

主持
6. 基于溶解氧氧同位素来揭示长江口外低氧区的溶解氧最低值。2026-2029.国家自然科学基金委员会 面上基金项目

5. 基于溶解氧氧同位素来量化水柱和沉积物耗氧对低氧贡献的演替特征。2020-2023. 国家自然科学基金委员会 面上基金项目
4. 冰川融水对北极近海峡湾中惰性溶解有机碳积累的影响。2017-2020. 国家自然科学基金委员会 面上基金项目
3. 贫营养/富营养河口中有机质的生物地球化学行为对比：以脂类和氨基酸对映体为例 2014-2015 科技部 中国-克罗地亚双边政府间科技合作项目
2. A study on dissolved organic nitrogen in the Changjiang Estuary: begin with amino acids enantiomers. 2012-2014. International Foundation for Science, Stockholm, Sweden. 【国际科学基金（瑞典）；类似国内面上项目】
1. 低纬度小河流流域盆地新鲜有机质入海定量化研究. 2013-2015. 国家自然科学基金委员会 青年基金项目

另外先后参加多项973或国家重点研发计划，及自然基金委重大项目

现场（野外）调查
 2011年5月东海陆架调查（973项目）船上首席科学家
 先后参加超过30次野外工作（包括大洋、近海河口、流域盆地、两极），野外总时间超过500天

Selected publications

1.      Fang, F., Zhu Z-Y*, Yi, Y., He, D., Bao, H., Zhang, E. and Zhou, C., 2025. Eutrophication-induced dinoflagellate succession contributes to marine carbon sequestration through refractory dissolved organic matter accumulation. Limnology and Oceanography, 70: 1992-2001.

2.      Zhu Z-Y*, Song, G.-D., Zhang, Z.-H., Luo, Y.-S., Liu, S.-M. and Zhang, J., 2025. On the stoichiometry of degradable fraction of particulate organic matter along a diapycnal mixing path using dissolved oxygen δ18O approach. Journal of Geophysical Research: Oceans, 130(6): e2025JC022512.

3.      Zhu, Z.-Y.*, 2022. Clarifying the fate of dissolved organic carbon in turbid glacier meltwater rivers in Svalbard via a series of incubations. Biogeochemistry. 159: 337-352

4.      Zhu, Z.-Y.*, Zhang, J., Wu, Y., Zhang, Y.-Y., Lin, J. and Liu, S.-M., 2011. Hypoxia off the Changjiang (Yangtze River) Estuary: Oxygen depletion and organic matter decomposition. Marine Chemistry, 125(1-4): 108-116.

5.      Zhuo Yi. Zhu*, Jing Zhang, Ying Wu and Jing Lin. Bulk particulate organic carbon in the East China Sea: Tidal influence and bottom transport. 2006. Progress in Oceanography. 69: 37-60.

完整论文清单（第一/通讯作者；截止2026年10月）:

1.      Zhang, Z.-H., Luo, Y.-S., Jin, J. and Zhu Z-Y*, 2026. Optimization of Gas Chromatography-Isotope Ratio Mass Spectrometry Oxygen Isotope Analysis Method and Recommendations for Precision Control. Journal of Chromatography A, 1765: 466544.

2.      Hossen, S., FANG, F.-T., Yang, L.-Y., Zhang, Z.-H., Hao, Y.-Y., Uddin, S.A., Lin, Z.-Z., Zhou, C.-X. and Zhu Z-Y*, 2026. Resuspension-driven organic matter dynamics in bivalve aquaculture systems: seasonal insights from the Dinghai Bay, China. Aquaculture Environment Interactions, 18: aei00507.

3.      Zhou, Y., Yang, L.*, Fang, F., Li, Y., Zhuang, W.-E. and Zhu Z-Y*, 2026. Unveiling the molecular composition of dissolved organic phosphorus in the freshwater-estuarine continuum using ultrahigh-resolution FT-ICR-MS. Environmental Research, 289: 123352.

4.      Fang, F.-T., Zhu Z-Y*, Yi, Y.-B. and He, D., 2025. Seasonality of Surface Dissolved Organic Matter Composition in the Northern Part of a Large Marginal Sea in an Extremely Dry Year. Journal of Geophysical Research: Oceans, 130(10): e2024JC022019.

5.      Fang, F., Zhu Z-Y*, Yi, Y., He, D., Bao, H., Zhang, E. and Zhou, C., 2025. Eutrophication-induced dinoflagellate succession contributes to marine carbon sequestration through refractory dissolved organic matter accumulation. Limnology and Oceanography, 70: 1992-2001.

6.      Zhang, Z.-H., Zhu Z-Y*, Zhang, W.-X., Xiao, C.-B., Fan, W. and Tang, K., 2025. Sedimentary and water column contributions to oxygen depletion considering mixing in a dynamic coast. Estuarine, Coastal and Shelf Science, 320: 109300.

7.      Zhu Z-Y*, Song, G.-D., Zhang, Z.-H., Luo, Y.-S., Liu, S.-M. and Zhang, J., 2025. On the stoichiometry of degradable fraction of particulate organic matter along a diapycnal mixing path using dissolved oxygen δ18O approach. Journal of Geophysical Research: Oceans, 130(6): e2025JC022512.

8.      Huang, H.*, Li, Y., Zhu Z-Y*, Cheng, P.-F., Zhao, X. and Dang, S.-N., 2025. Study on carbon budget in small-scale high-density intensive Litopenaeus vannamei aquaculture ponds. Aquaculture International, 33(6): 491.

9.      Li, Y., Huang, H.*, Zhu Z-Y*, Cheng, P., Zhao, X. and Dang, S.-N., 2025. Quantifying and characterizing degradation of dissolved organic carbon from aquaculture wastewater: a case study of Litopenaeus vannamei. Frontiers in Marine Science, 12: 1627724.

10.   Fang, F.-T., Zhu Z-Y*, Wenger, F., Ge, J.-Z., Du, J.-Z., Deng, B., Ma, H.-M., Zhang, R.-F. and Zhang, Y., 2024. Short-term sedimentary evidence for increasing diatoms in Arctic fjords in a warming world. Science of The Total Environment, 951: 175757.

11.   He, J.-Y.*, Tao, Y.-L., Shao, S., Wei, H., Yan, G.-X., Tang, C.-Y., Feng, J., Li, M.-S., Liao, Z., Zhang, X.-L., Tang, C.-S., Buttino, I., Wang, J.-X., Zhu Z-Y* and Yan, X.-J.*, 2024. The hidden acceleration pump uncovers the role of shellfish in oceanic carbon sequestration. Science of The Total Environment, 951: 175699.

12.   Li, Y.-Q., Fang, F.-T., Zeng, C. and Zhu Z-Y*, 2024. Dissolved oxygen fluctuation alters dissolved organic carbon stability and can accumulate resistant fraction. Marine Chemistry: 104441.

13.   Zhang, W., Wang, Y., Fang, F., Zhuang, W.-E., Yang, L.* and Zhu Z-Y*, 2024. Characterization of sediment organic matter in the outer Yangtze River Estuary using stable isotopes, optical techniques, and FT-ICR-MS: Implications for the carbon burial mechanism. Water Research: 122512.

14.   Zhang, Z.-H., Zhu Z-Y*, Hu, H.-T., Zhou, J., Lin, Z.-Z. and Zhou, C.-X., 2024. Quantifying sedimentary oxygen consumption contribution to total oxygen depletion in shallow waters via the oxygen isotope approach. Marine Ecology Progress Series, 728: 1-14.

15.   Zhu Z-Y*, Fang F-T, Hossen S, Zhao Y, Chen Z-Z, Zeng C, Zhou M (2023) Impact of wild fish faeces to deep sea dissolved organic carbon as revealed by laboratory incubations. Deep Sea Research Part I: Oceanographic Research Papers 200: 104148.

16.   Jiakang Li, Zhuoyi Zhu*, Zhifeng Yang, Weiyi Li, Yongxin Lv, Yu Zhang (2023) Soil microorganisms and methane emissions in response to short-term warming field incubation in Svalbard. Frontiers in Microbiology 14: 1276065

17.   Cao Y, Huang H*, Zhao X, Li J, Wang T, Lin J, Zhu Z-Y* (2023) Labile and refractory fractions of sedimentary organic carbon off the Changjiang Estuary and its implications for sedimentary oxygen consumption. Frontiers in Marine Science 9: 1096108

18.   Zhu, Z.-Y.*, 2022. Clarifying the fate of dissolved organic carbon in turbid glacier meltwater rivers in Svalbard via a series of incubations. Biogeochemistry. 159: 337-352

19.   Zhou, M., Fang, F.-T., Zeng, C., Zhang, L.-H., Zhou, C.-X., Zhu, Z.-Y.*, 2022. Community competition is the microorganism feedback to sedimentary carbon degradation process in aquaculture tidal flats. Frontiers in Marine Science, 9: 880120.

20.   Fang, F.-T., Zhu, Z.-Y.*, Ge, J.-Z., Deng, B., Du, J.-Z., Zhang, J., 2022. Reconstruction of the main phytoplankton population off the Changjiang Estuary in the East China Sea and its assemblage shift in recent decades: From observations to simulation. Marine Pollution Bulletin 178, 113638.

21.   Yang L*, Chen Y, Lei J, Zhu Z* (2022) Effects of coastal aquaculture on sediment organic matter: Assessed with multiple spectral and isotopic indices. Water Research: 118951

22.   Zhu, Z.Y.*, Wang, J., Zhang, G.L., Liu, S.M., Zheng, S., Sun, X.X., Xu, D.F., Zhou, M., 2021. Using triple oxygen isotopes and oxygen-argon ratio to quantify ecosystem production in the mixed layer of northern South China Sea slope region. Acta Oceanologica Sinica 40, 1-15.

23.   Zhou, J., Zhu, Z.Y.*, Hu, H.T.*, Zhang, G.L., Wang, Q.Q., （2021）Clarifying water column respiration and sedimentary oxygen respiration under oxygen depletion off the Changjiang Estuary and adjacent East China Sea. Frontiers in Marine Science. 7，623581.

24.   Hao Y-Y, Zhu Z-Y*, Fang F-T, Novak T, Canković M, Hrustić E, Ljubešić Z, Li M, Du J-Z, Zhang R-F, Gašparović B (2021) Tracing Nutrients and Organic Matter Changes in Eutrophic Wenchang (China) and Oligotrophic Krka (Croatia) Estuaries: A Comparative Study. Frontiers in Marine Science 8(1141): 663601

25.   Lei, J., Yang, L.*, and Zhu, Z.* (2021). Testing the effects of coastal culture on particulate organic matter using absorption and fluorescence spectroscopy. Journal of Cleaner Production 325, 129203.

26.   Zhu, Z.Y.*, Oakes J, Eyre B, Hao Y, Sia ESA, Jiang S, Müller M, Zhang J (2019) The non-conservative distribution pattern of organic matter in Rajang, a tropical river with peatland in its estuary. Biogeosciences. 17, 2473-2485. doi:10.5194/bg-2019-157

27.   Zhu, Z.Y.*, Wu, H., Liu, S.M., Wu, Y., Huang, D.J., Zhang, j. and Zhang, G.S., 2017. Hypoxia off the Changjiang (Yangtze River) Estuary and in the adjacent East China Sea: quantitative approaches to estimating the tidal impact and nutrient regeneration. Marine Pollution Bulletin, 125(1-2): 103-114.

28.   Zhu, Z.Y.*, J. Hu, G.D. Song, Y. Wu, J. Zhang, and S.M. Liu, Phytoplankton-driven dark plankton respiration in the hypoxic zone off the Changjiang Estuary, revealed by in vitro incubations. Journal of Marine Systems, 2016. 154, Part A: p. 50-56.

29.   Zhu, Z.Y.*, Y. Wu, S.M. Liu, F. Wenger, J. Hu, J. Zhang, and R.F. Zhang, Particulate organic matter composition and organic carbon flux in Arctic valley glaciers: examples from the Bayelva River and adjacent Kongsfjorden. Biogeosciences, 2016. 13(18): p. 975-987.

30.   Zhu, Z.Y.*, S.M. Liu, Y. Wu, Y. Li, J. Zhang, and J. Hu, Phytoplankton Dynamics and Its Further Implication for Particulate Organic Carbon in Surface Waters of a Tropical/Subtropical Estuary. Estuaries and Coasts, 2015. 38(3): p. 905-916.

31.   Zhu, Z. Y.*, Y. Wu, J. Zhang, J. Z. Du, and G. S. Zhang (2014), Reconstruction of anthropogenic eutrophication in the region off the Changjiang Estuary and central Yellow Sea: From decades to centuries, Continental Shelf Research, 72, 152-162.

32.   Zhu, Z. Y.*, Y. Wu, J. Zhang, T. Dittmar, Y. Li, L. Shao, and Q. Ji (2014), Can primary production contribute non-labile organic matter in the sea: Amino acid enantiomers along the coast south of the Changjiang Estuary in May, Journal of Marine Systems, 129, 343-349.

33.   Zhu, Z.-Y.*, Zhang, J., Wu, Y., Zhang, Y.-Y., Lin, J. and Liu, S.-M., 2011. Hypoxia off the Changjiang (Yangtze River) Estuary: Oxygen depletion and organic matter decomposition. Marine Chemistry, 125(1-4): 108-116.

34.   Zhu, Z.Y.*, Ng, W.M., Liu, S.M., Zhang, J., Chen, J.C. and Wu, Y., 2009. Estuarine phytoplankton dynamics and shift of limiting factors: A study in the Changjiang (Yangtze River) Estuary and adjacent area. Estuarine, Coastal and Shelf Science, 84: 393-401.

35.   Zhuo Yi. Zhu*, Jing Zhang, Ying Wu and Jing Lin. Bulk particulate organic carbon in the East China Sea: Tidal influence and bottom transport. 2006. Progress in Oceanography. 69: 37-60.

36.   Zhu Zhuoyi*, Wu Ying, Zhang Jing, Wei Hao. Spatial and temporal variation of particulate organic carbon in the PN section of East China Sea. 2005. Acta Oceanologica Sinica. 24(5):89-99.

学术论著（章节）
Zhang, J., Wu, Y., Zhang, G.L. and Zhu, Z.Y., 2014. Carbon biogeochemistry in the continuum of the Changjiang (Yangtze) River watersheds across the East China Sea. In: T. Bianchi, M. Allison and W.J. Cai (Editors), Biogeochemical dynamics at major river-coastal interfaces: Linkages with global change. Cambridge University Press, New York, pp. 237-273.
Zhu Z (2021) Hypoxia. In: Ishizaka J, Kim G, Lee JH, Liu SM, Yu F & Zhang J (eds) Oceanography of the Yellow Sea and East China Sea. PICES Science Report No. 62. North Parcific Marine Science Organization, Sidney, Canada. p 121-137

其他合作者身份论文略

海洋化学与AI（本科）

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微波消解仪的转子内衬装置；朱卓毅 李月；ZL2022 2 1783073.9
一种采集中底层水的铅鱼配重装置; 朱卓毅；ZL 2022 2 3367824.1

中国海洋湖沼学会化学分会 理事，2022-
中国海洋学会海洋化学分会 委员，2022-
Frontiers in Marine Science 编辑
自然资源部大亚湾海洋风险与灾害野外科学观测研究站 副站长

期刊评审：
Geophysical Research Letters
Limnology and Oceanography
The Cryosphere
Science of the Total Environment
Journal of Geophysical Research: Oceans
Journal of Geophysical Research: Biogeoscience
Biogeosciences
Marine Chemistry
Biogeochemistry

1999 刘翔奖学金
2007 上海市优秀毕业生